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The free tripeptide chelates a single Cu(II) ion through its N-terminal amine, the deprotonated glycyl amide nitrogen and the histidine imidazole, giving the square-planar complex responsible for the deep blue colour of a reconstituted ghkcu solution

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Xiong H, Wang J, Ran Q, Lou G, Peng C, Gan Q, Hu J, Sun J, Yao R, Huang Q (2019) Hesperidin: a therapeutic agent for obesity

The role of CLASPs and LL5 in integrin-regulated microtubule stabilization is discussed in more detail below